Articles published in scientific journals


  1. Blaimer, B., Santos, B.F., Cruaud, A., Gates, M.W., Kula, R.R., Mikó, I., Rasplus, J-Y., Smith, D.R., Talamas, E.J., Brady, S.G. & Buffington, M.L. 2023. Key innovations and the diversification of Hymenoptera. Nature Communications, 14: 1212.
  2. Santos, B.F., Miller, M.E., Miklasevskaja, M., McKeown, J.T.A., Redmond, N.E., Coddington, J.A., Bird, J., Miller, S.E., Smith, A., Brady, S.G., Buffington, M.L., Chamorro, M.L., Dikow, T., Gates, M.W., Goldstein, P., Konstantinov, A., Kula, R., Silverson, N.D., Solis, M.A., deWaard, S.L., Naik, S., Nikolova, N., Pentinsaari, M., Prosser, S.W.J., Sones, J.E., Zakharov, E.V., deWaard, J.R. 2023. Enhancing DNA barcode reference libraries by harvesting terrestrial arthropods at the Smithsonian`s National Museum of Natural History. Biodiversity Data Journal, 11: e100904.
  3. Neves, K., Santos, B.F., Schultz, T., Gotzek, D., Abreu, R.C.R., Durigan, G. &  Vasconcelos, H.L. 2023. Woody encroachment affects multiple dimensions of ant diversity in a neotropical savanna. Insect Conservation and Diversity.
  4. Supeleto, F.A., Santos, B.F. & Aguiar, A.P. 2022. Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa, 5219:501-533.
  5. Santos, B.F., Sandoval, M., Spasojevic, T., Giannotta, M.M. & Brady, S.G. 2022. A Parasitoid Puzzle: Phylogenomics, Total-evidence Dating, and the Role of Gondwanan Vicariance in the Diversification of Labeninae (Hymenoptera, Ichneumonidae). Insect Systematics and Diversity, 6(5): 1-13. DOI: 10.1093/isd/ixac015 [PDF]

  6. Jasso-Martínez, J.M.*, Santos, B.F.*, Zaldívar-Riverón, A., Fernández-Triana, J.L., Sharanowski, B.J., Richter, R., Dettman, J.R., Blaimer, B.B. Brady, S.G. & Kula, R.R. 2022. Phylogenomics of braconid wasps (Hymenoptera, Braconidae) sheds light on classification and the evolution of parasitoid life history traits. Molecular Phylogenetics and Evolution, 173:107452. [PDF] (*co-first authors)
  7. Jasso-Martínez, J.M., Quicke, D.L.J., Belokobylskij, S.A., Santos, B.F., Fernández-Triana, J.L., Kula, R.R. & Zaldívar-Riverón, A. 2022. Mitochondrial phylogenomics and mitogenome organization in the parasitoid wasp family Braconidae (Hymenoptera: Ichneumonoidea). BMC Ecology and Evolution, 22:46, [PDF]

  8. Pádua, D.G., Kloss, T.G., Tavares, M.T., Santos, B.F., Araújo, R.O., Schoeninger, K., Sobczak, J.F. & Gonzaga, M.O. 2022. Hyperparasitoids of polysphinctine Darwin wasps (Hymenoptera: Ichneumonidae) in South America. Austral Entomology, 61:170-186.

  9. Bordera, S. & Santos, B.F. 2022. A review of Piasites Seyrig (Hymenoptera, Ichneumonidae, Cryptinae), with description of seven new species. Journal of Hymenoptera Research, 90:23–57.[PDF]

  10. Santos, B.F.*, Klopfstein, S.*, Whitfield, J.B.* & Sharanowski, B.J.* 2022. Many evolutionary roads led to virus domestication in ichneumonoid parasitoid wasps. Current Opinion in Insect Science, 50:100861. [PDF] (*co-first authors)

  11.  Samacá-Sáenz, E., Santos, B.F., Martínez, J.J., Egan, S.P., Shaw, S.R., Hanson, P.E. & Zaldívar-Riverón, A. 2022. Ultraconserved elements-based systematics reveals evolutionary patterns of host-plant family shifts and phytophagy within the predominantly parasitoid braconid wasp subfamily Doryctinae. Molecular Phylogenetics and Evolution, 166: 107319. [PDF]

  12. Santos, B.F., Wahl, D.B., Rousse, P., Bennett, A.M.R., Kula, R. & Brady, S.G. 2021. Phylogenomics of Ichneumoninae (Hymenoptera, Ichneumonidae) reveals pervasive morphological convergence and the shortcomings of previous classifications. Systematic Entomology, 46: 704–724. [PDF]

  13. Supeleto, F.A., Aguiar, A.P. & Santos, B.F. 2021. A new species, key and further redefinition of Nesolinoceras Ashmead (Hymenoptera, Ichneumonidae, Cryptinae). Zootaxa, 5016: 107–116.[PDF]


  14. Somavilla, A., Santos, B.F., Carpenter, J.M., Andrena, S.R. & Oliveira, M.L. 2021. Total-evidence phylogeny of the New World Polistes Lepeletier, 1836, paper wasps (Vespidae, Polistinae, Polistini). American Museum Novitates, 3973: 1–42. [PDF]

  15. Sandoval, M. & Santos, B.F. 2021. A new species of Grotea Cresson, the first record of Labeninae (Hymenoptera, Ichneumonidae) in the Greater Antilles. Journal of Hymenoptera Research, 81: 1–8. [PDF]


  16. Legeai, F*., Santos, B.F.*, Robin, S.*, Breteaudeau, A., Dikow, R., Lemaitre, C., Jouan, V., Ravallec, M. Drezen, J.M., Tagu, D., Gyapay, G., Zhou, X., Liu, S., Webb, B., Brady, S.G. & Volkoff, A.N. 2020. Genomic architecture of endogenous ichnoviruses reveals distinct evolutionary pathways leading to virus domestication in parasitic wasps. BMC Biology, 18:89. [PDF] (*co-first authors)

  17. Supeleto, F.A., Santos, B.F., Basílio, L.A. & Aguiar, A.P. 2020. Species delimitation, environmental cline and phylogeny for a new Neotropical genus of Cryptinae (Ichneumonidae). PLoS ONE, 15(10): e0237233. DOI: 10.1371/journal.pone.0237233. [PDF]

  18. Supeleto, F.A., Santos, B.F. & Aguiar, A.P. 2020. A new species and southernmost record of Cestrus Townes (Hymenoptera, Ichneumonidae, Cryptinae). Zootaxa, 4822: 277–284 [PDF]

  19. Supeleto, F.A., Santos, B.F. & Aguiar, A.P. 2020. Phylogenomic analyses reveal a rare new genus of wasp (Hymenoptera, Ichneumonidae, Cryptinae) from the Brazilian Atlantic Forest. Systematics and Biodiversity, 1–16. [PDF]

  20. Klopfstein, S., Santos, B.F., Shaw, M.R., Alvarado, M., Bennett, A.M., Dal Pos, D., Giannotta, M., Herrera-Florez, A.F., Karlsson, D., Khalaim, A.I., Lima, A.R., Mikó, I., Sääksjärvi, I. E., Shimizu, S., Spasojevic, T., van Noort, S., Vilhelmsen, L., & Broad, G. R. 2019. Darwin wasps: a new name heralds renewed efforts to unravel the evolutionary history of Ichneumonidae. Entomological Communications, 1: ec01006. DOI:10.37486/2675-1305.ec01006 [PDF]

  21. Reshchikov, A., Santos, B.F., Liu, J.X. & Barthélémy, C. 2019. Review of Palpostilpnus Aubert (Hymenoptera, Ichneumonidae, Phygadeuontinae), with the description of ten new species. European Journal of Taxonomy, 582: 1–63. DOI: 10.5852/ejt.2019.582 [PDF]

  22. Supeleto, F.A., Santos, B.F. & Aguiar, A.P. 2019. Revision of Distictus Townes, 1966 (Hymenoptera, Ichneumonidae, Cryptinae), with descriptions of ten new species. European Journal of Taxonomy, 542: 1–64. DOI: 10.5852/ejt.2019.542. [PDF]

  23. Santos, B.F., Perrard, A. & Brady, S.G. 2019. Running in circles in phylomorphospace: host environment constrains morphological diversification in parasitic wasps. Proceedings of the Royal Society B: Biological Sciences, 286:20182352. [PDF]

  24. Santos, B.F., Alvarado, M., Sääksjärvi, I.E., van Noort, S., Villemant, C. & Brady, S.G. 2018. Molecular phylogeny of Ateleutinae (Hymenoptera: Ichneumonidae): systematics and biogeography of a widespread parasitoid wasp lineage. Zoological Journal of the Linnean Society, 1–22. DOI: 10.1093/zoolinnean/zly072. [PDF]

  25. Araújo, R.O., Vivallo, F. & Santos, B.F. 2018. Ichneumonid wasps of the subfamily Mesochorinae: new replacement names, combinations and an updated key to the World genera (Hymenoptera: Ichneumonidae). Zootaxa, 4521:52–60. DOI: 10.11646/zootaxa.4521.1.2. [PDF]

  26. Santos, B.F. & Perrard, A. 2018. Testing the Dutilleul syndrome: host use drives the convergent evolution of multiple traits in parasitic wasps. Journal of Evolutionary Biology,  31:1430–1439. DOI: 10.1111/jeb.13343. [PDF]

  27. Santos, B.F. & Hoppe, J.P.M. 2018. Filling gaps in species distributions through the study of biological collections: 415 new distribution records for Neotropical Cryptinae (Hymenoptera, Ichneumonidae). Revista Brasileira de Entomologia, 62:288–291. DOI: 10.1016/j.rbe.2018.09.001. [PDF]

  28. Santos, B.F., Scherrer, M.V. & Loss, A.C. 2018. Neither barriers nor refugia explain genetic structure in a major biogeographic break: phylogeography of praying mantises in the Brazilian Atlantic Forest. Mitochondrial DNA Part A, 29:1284–1292 DOI: 10.1080/24701394.2018.1445242. [PDF]

  29. Santos, B.F. & Aguiar, A.P. 2018. Review of Dotocryptus Brèthes (Hymenoptera, Ichneumonidae, Cryptinae), with a New Species from Colombia. Neotropical Entomology, 47:871–844. DOI 10.1007/s13744-018-0602-y. [PDF]

  30. Araújo, R.O., Vivallo, F. & Santos, B.F. 2018. Discovery of two new Andean species of Scolomus (Townes), with a key to all known species (Hymenoptera: Ichneumonidae: Metopiinae). Zootaxa, 4429: 189–194. DOI: 10.11646/zootaxa.4429.1.12 [PDF]

  31. Santos, B.F. 2017. Phylogeny and reclassification of Cryptini (Hymenoptera, Ichneumonidae, Cryptinae), with implications for ichneumonid higher-level classification. Systematic Entomology, 42:650–676. DOI 10.1111/syen.12238. [PDF]

  32. Santos, B.F., Aguiar, A.P., Tedesco, A.M. & Fontenelle, J.C.R. 2017. Long-term seasonal dominance of the wasp Trihapsis polita Townes (Hymenoptera, Ichneumonidae) in the Brazilian Atlantic Forest. Biodiversity Data Journal, 5, e11337. [PDF]

  33. Santos, B.F. 2016. Generic redefinition and a new species of Nesolinoceras (Hymenoptera, Ichneumonidae, Cryptinae). American Museum Novitates, 3858: 1–16. [PDF]

  34. Villanueva-Bonilla, G.A., Onody, H.C., Santos, B.F. & Vasconcelos-Neto, J. 2016. First record of egg sac predation on a wall crab spider Selenopidae (Araneae) by the wasp Camera lunavenatrix n. (Ichneumonidae, Cryptinae). Journal of Hymenoptera Research, 49: 65–79. [PDF]

  35. Aguiar, A.P. & Santos, B.F. Revision of Melanocryptus Cameron (Ichneumonidae, Cryptinae), with description of seven new species. American Museum Novitates, 3836: 1–56. [PDF]

  36. Santos, B.F., Payne, A., Pickett, K.M. & Carpenter, J.M. 2015. Phylogeny and historical biogeography of the paper wasp genus Polistes (Hymenoptera: Vespidae): implications for the overwintering hypothesis of social evolution. Cladistics, 31: 535–549. [PDF]

  37. Santos, B.F. & Aguiar, A.P. 2015. Review of Loxopus Townes (Hymenoptera, Ichneumonidae, Cryptinae), with descriptions of six new species. Journal of Natural History, 49(31-32): 1905–1935. [PDF]

  38. Santos, B.F. & Aguiar, A.P. 2013. Phylogeny and revision of Messatoporus Cushman (Hymenoptera, Ichneumonidae, Cryptinae), with description of sixty five new species. Zootaxa 3634: 1–284. [PDF]

  39. Aguiar, A.P. & Santos, B.F. 2012. Additions to the revision of Digonocryptus Viereck (Hymenoptera, Ichneumonidae, Cryptinae), with nine new taxa, new males and distribution maps for all known species. Zootaxa, 3828: 1–41. [PDF]

  40. Santos, B.F., Aguiar, A.P. & Tedesco, AM. 2012. Trigonalidae (Hymenoptera) from cacao agroforestry systems in northeastern Brazil, with two new species of Trigonalys Journal of Hymenoptera Research, 25: 19–33. [PDF]

  41. Santos, B.F. & Aguiar, A.P. 2012. Phylogeny and description of Eknomia, a morphologically unusual new genus of Neotropical Cryptinae (Hymenoptera, Ichneumonidae, Cryptinae). Zootaxa, 3237: 35–52. [PDF]

  42. Aguiar, A.P. & Santos, B.F. 2010. Discovery of potent, unsuspected sampling disparities for Malaise and Möricke traps, as shown for Neotropical Cryptini (Hymenoptera, Ichneumonidae). Journal of Insect Conservation, 14: 199–2010. [PDF]

  43. Scherrer, M.V., Santos, B.F. & Aguiar, A.P. 2010. Generic redefinition, key and two new species of Acorystus Townes (Hymenoptera, Ichneumonidae, Cryptinae). Zootaxa, 2721: 28–38. [PDF]

  44. Santos, B.F., Aguiar, A.P. & Tedesco, A.M. 2009. Phylogenetic revision and the origin of Polyphrix Townes (Hymenoptera, Ichneumonidae, Cryptinae), with description of a new species. Zootaxa, 1934: 30–39. [PDF]

  45. Aguiar, A.P. & Santos, B.F. 2009. Biological synopsis of Photocryptus Viereck (Hymenoptera, Ichneumonidae, Cryptinae), with eight new host records and fourteen new geographic records. Zootaxa, 2148: 49–45. [PDF]

  46. Santos, B.F. & Aguiar, A.P. 2008. Phylogeny and reclassification of Distictus Townes (Hymenoptera, Ichneumonidae, Cryptinae), with description of a new species. Zootaxa, 1934: 30–39. [PDF]

Book chapter

Aguiar, A.P., Santos, B.F., Couri, M.S., Rafael, J.A., Costa, C. Ide, S., Duarte, M., Grazia, J., Schwertner, C.F., Freitas, A.V.L. & Azevedo, C.O. Insecta. In: Rocha, R.M. & Boeger, W.A. Estado da Arte e Perspectivas para a Zoologia no Brasil [State of Art and Perspectives to the Zoology in Brazil]. Editora UFPR, Curitiba, pp.131–155. [PDF in Portuguese]